Classification / Names
Common names | Synonyms | Catalog of Fishes (gen., sp.) | ITIS | CoL | WoRMS | Cloffa
Actinopterygii (ray-finned fishes) > Perciformes
(Perch-likes) > Gobiidae
(Gobies) > Gobiinae
Etymology: papilio: Name from Latin 'papilio' meaning butterfly, used as the generic name of the swallowtail butterfly, refers to the elongate, filamentous fifth and ninth branched caudal-fin rays (like the 'tails' of the swallowtail butterfly); noun in apposition.
Environment: milieu / climate zone / depth range / distribution range
Marine; demersal; depth range 5 - 55 m (Ref. 90102). Temperate
Northwest Pacific: Ryukyu Is., Japan.
Size / Weight / Age
Maturity: Lm ?  range ? - ? cm
Max length : 4.1 cm SL male/unsexed; (Ref. 55718)
Morphology | Morphometrics
soft rays: 11;
Vertebrae: 26. This species is distinguished from its congeners in having the following set of characters: segmented dorsal-fin rays 11; segmented anal-fin rays 11; longitudinal scales 27; predorsal scales 11; third spine of first dorsal fin elongate, filamentous, and longer than the other spines; fifth and ninth branched caudal-fin rays elongate, filamentous, and forming bifurcate caudal fin; fifth and ninth branched rays each with a single branch; cephalic sensory-papillae row 'a' with 3 widely spaced papillae; numerous small yellow spots scattered on cheek, operculum, nape, body, and dorsal fins in fresh or live individuals; below the eye is a dusky yellowish brown vertical bar; body with 4 dark gray brown saddles, in addition to a similarly colored large blotch at midlateral caudal-fin base (Ref. 55718).
A single specimen collected from a sandy-mud bottom at a depth of 45 m in a protected bay. Observed to be usually at 40-50 m depths (area with pebbles); the young at depths of 8 m to about 30 m, but apparently on rare occasions; the species symbiotically associates with alpheid shrimp (species name uncertain) (Ref. 55718).
Life cycle and mating behavior
Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae
Shibukawa, K. and T. Suzuki, 2004. Vanderhorstia papilio, a new shrimp-associated goby from the Ryukyu Islands, Japan (Perciformes: Gobiidae: Gobiinae), with comments on the limits of the genus. Ichthyol. Res. 51:113-119. (Ref. 55718)
IUCN Red List Status (Ref. 119314)
CITES (Ref. 115941)
Threat to humans
Common namesSynonymsMetabolismPredatorsEcotoxicologyReproductionMaturitySpawningSpawning aggregationFecundityEggsEgg development
ReferencesAquacultureAquaculture profileStrainsGeneticsAllele frequenciesHeritabilityDiseasesProcessingMass conversion
Estimates of some properties based on models
Preferred temperature (Ref. 115969
): 23.6 - 28.9, mean 27.9 (based on 202 cells).
Phylogenetic diversity index (Ref. 82805
= 0.5000 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00661 (0.00300 - 0.01455), b=3.08 (2.90 - 3.26), in cm Total Length, based on LWR estimates for this (Sub)family-body shape (Ref. 93245
Trophic Level (Ref. 69278
): 3.2 ±0.3 se; Based on size and trophs of closest relatives
Resilience (Ref. 69278
): High, minimum population doubling time less than 15 months (Preliminary K or Fecundity.).
Vulnerability (Ref. 59153
): Low vulnerability (10 of 100) .